Research Group Leader: Dr Richard J. Butler
School of Geography, Earth and Environmental Sciences
University of Birmingham
Edgbaston, Birmingham, B15 2TT
butler.richard.j
[at]gmail.com
r.butler.1[at]bham.ac.uk



I am a vertebrate palaeontologist whose research focuses on the systematics, evolution and biogeography of late Palaeozoic to Mesozoic reptiles. I am particularly interested in the origin and dramatic evolutionary radiation of dinosaurs and closely related fossil groups in the aftermath of the Permo-Triassic mass extinction, the largest extinction event in the history of life on Earth. Another major research focus addresses the patterns and drivers of vertebrate diversity and body size over extended geological timescales. 

I have published >75 peer-reviewed papers, established a highly active network of cooperative projects worldwide, and my work has been recognised by awards from professional societies in the UK and USA. My research is currently funded by the Emmy Noether Program of the German Research Foundation and a Career Integration Grant from the Marie Curie Actions.


POSITIONS AND EDUCATION

Current

Since 2013–. Birmingham Fellow (permanent, research-intensive appointment at the Lecturer level) and Academic Keeper of the Lapworth Museum of Geology, School of Geography, Earth and Environmental Sciences, University of Birmingham, UK.

2014–2016. Honorary Senior Research Fellow, Evolutionary Studies Institute, University of the Witwatersrand, Johannesburg, South Africa. 

Past

2011–2013. Junior Research Group Leader, Emmy Noether Programme (DFG), GeoBio-Center, LMU, Munich, Germany. 

2009–2011. Alexander von Humboldt Foundation Research Fellowship. Bayerische Staatssammlung für Paläontologie und Geologie, Munich, Germany

2008–2009. NERC Researcher Co-Investigator. Natural History Museum London.

2006–2008. NERC Postdoctoral Research Assistant. Natural History Museum, London.

2002–2007. Ph.D. (NERC-CASE funded). University of Cambridge and Natural History Museum, London.

1999–2002. BSc. Geology (First Class Honours). Department of Earth Sciences, University of Bristol.


KEY GRANT FUNDING (Full details on request)

2014–2018. Marie Curie Career Integration Grant: "The early Mesozoic rise of archosaurs: New insights into an exemplar evolutionary radiation" (Principal Investigator)

2011–2014. German Research Foundation (DFG) Emmy Noether Programme: “Dawn of the dinosaurs: Archosauromorph evolution in the terrestrial Triassic” (Principal Investigator)

2010–2011. German Research Foundation (DFG) Research Grant: “Terrestrial vertebrates from near the Triassic-Jurassic boundary in Portugal: Excavation, geological context, and faunal change” (
Principal Investigator)

2008–2009. NERC Small Grant: “Origin of the avian respiratory system: A CT-study of postcranial pneumaticity in basal archosaurs” (Researcher Co-Investigator)

Plus 16 additional small grants (each < £5000) from
Royal Society, Synthesys, Lamont-Doherty Earth Observatory, Jurassic Foundation, Palaeontological Association, University of Cambridge, Systematics Association, Cambridge Philosophical Society, Samuel & Doris Welles Research Fund and Percy Sladen Memorial Fund.


AWARDS

2011. Hodson Award, Palaeontological Association (link).

2005. Predoctoral Fellowship, Society of Vertebrate Paleontology (link). 
 


EDITORIAL ROLES

2013–. Editorial Board, Journal of Vertebrate Paleontology.

2013–. Editorial Board, Acta Palaeontologica Polonica.

2012–2014. Editorial Board, PLOS ONE.

2010–2013. Associate Editor, Paläontologische Zeitschrift.

2008–2011. Section Editor, Fossil Reptiles, Zootaxa.


COMMITTEE MEMBERSHIPS

2014–. Ordinary Member of Council, the Palaeontological Association.

2013. Executive Committee, Paleobiology Database.

2012–. Program Committee, Society of Vertebrate Paleontology.



CURRENT UNDERGRADUATE TEACHING

Module lead for ESCM 431, Advanced Projects
Three lectures on ESCM319, Evolution of Vertebrates
Contributor to Field Skills 1 and 2
Supervise BSc/MSci research projects/advanced projects in Palaeobiology & Palaeoenvironments


PUBLICATIONS

Google Scholar citation details are here. My current Google Scholar H-index is 24, with >1600 citations. 
Visit my Academia.edu page for PDFs.


Edited special volumes

Mannion, P. D., Benson, R. B. J. & BUTLER, R. J., editors. 2013. Vertebrate Palaeobiodiversity Patterns and the Impact of Sampling Bias. Palaeogeography, Palaeoclimatology, Palaeoecology 372: 1-146.

BUTLER, R. J., Irmis, R. B. & Langer, M. C., editors. 2011 (for 2010). Late Triassic Terrestrial Biotas and the Rise of Dinosaurs. Earth and Environmental Science Transactions of the Royal Society of Edinburgh 101, parts 3 & 4: 189-426.


In press

80. Sookias, R. B., Sullivan, C., Liu, J. & BUTLER, R. J. Systematics of putative euparkeriids (Diapsida: Archosauriformes) from the Triassic of China. PeerJ.

79. Liu, J., BUTLER, R. J., Sullivan, C. & Ezcurra, M. D. Chasmatosaurus ultimus,’ a putative proterosuchid archosauriform from the Middle Triassic, is an indeterminate crown archosaur. Journal of Vertebrate Paleontology.

78. Brusatte, S. L., BUTLER, R. J., Mateus, O. & Steyer, J. S. A new species of Metoposaurus from the Late Triassic of Portugal and comments on the systematics and biogeography of metoposaurid temnospondyls. Journal of Vertebrate Paleontology.

77. Ezcurra, M. D. & BUTLER, R. J. (published online). Taxonomy of the proterosuchid archosauriforms (Diapsida: Archosauromorpha) from the earliest Triassic of South Africa, and implications for the early archosauriform radiation. Palaeontology.

76. Upchurch, P., Andres, B., BUTLER, R. J. & Barrett, P. M. (published online). An analysis of pterosaurian biogeography: implications for the evolutionary history and fossil record quality of the first flying vertebrates. Historical Biology.

75. Brusatte, S. L.*, BUTLER, R. J.*, Barrett, P. M., Carrano, M. T., Evans, D. C., Lloyd, G. T., Mannion, P. D., Norell, M. A., Peppe, D. J., Upchurch, P. & Williamson, T. E. (published online). The extinction of the dinosaurs. Biological Reviews.
*Joint lead authors; listed alphabetically

74. Sookias, R. B., Sennikov, A. G., Gower, D. J. & BUTLER, R. J. (published online). The monophyly of Euparkeriidae (Reptilia: Archosauriformes) and the origins of crown Archosauria: a revision of Dorosuchus neoetus from the Middle Triassic of Russia. Palaeontology.

73. Niedźwiedzki, G., Brusatte, S. L., Sulej, T. & BUTLER, R. J. (published online). Basal dinosauriform and theropod dinosaurs from the middle-late Norian (Late Triassic) of Poland: implications for Triassic dinosaur evolution and distribution. Palaeontology

2014

72. Gower, D. J., Hancox, P. J., Botha-Brink, J., Sennikov, A. G. & BUTLER, R. J. A new species of Garjainia Ochev, 1958 (Diapsida: Archosauriformes: Erythrosuchidae) from the Early Triassic of South Africa. PLoS ONE 9(11): e111154.

71. Barrett, P. M., BUTLER, R. J., Mundil, R., Scheyer, T. M., Irmis, R. B. & 
Sánchez-Villagra, M. R. 2014. A palaeoequatorial ornithischian and new constraints on early dinosaur diversification. Proceedings of the Royal Society B 281: 20141147 

70. Mateus, O., BUTLER, R. J., Brusatte, S. L., Steyer, J. S. & Whiteside, J. H. 2014. The first phytosaur (Diapsida, Archosauriformes) from the Late Triassic of the Iberian Peninsula. Journal of Vertebrate Paleontology 34:970-975.

69. BUTLER, R. J., Sullivan, C., Ezcurra, M. D., Liu, J., Lecuona, A. & 
Sookias, R. B. 2014. New clade of enigmatic early archosaurs yields insights into early pseudosuchian phylogeny and the biogeography of the archosaur radiation. BMC Evolutionary Biology 14:128. 

68. Benson, R. B. J., Frigot, R. A., Goswami, A., Andres, B. & BUTLER, R. J. 2014. Competition and constraint drove Cope's rule in the evolution of giant flying reptiles. Nature Communications 5:3567.

67. Ezcurra, M. D., Scheyer, T. & BUTLER, R. J. 2014. The origin and early evolution of Sauria: reassessing the Permian saurian fossil record and the timing of the crocodile-lizard divergence. PLOS ONE 9: e89165.

66. BUTLER, R. J., Rauhut, O. W. M., Stocker, M. R. & Bronowicz, R. 2014. Redescription of the phytosaurs Paleorhinus ("Francosuchus") angustifrons and Ebrachosuchus neukami from Germany, with implications for Late Triassic biochronology. Zoological Journal of the Linnean Society 170: 155–208

2013

65. Nesbitt, S. J., BUTLER, R. J. & Gower, D. J. 2013. A new archosauriform (Reptilia: Diapsida) from the Manda beds (Middle Triassic) of southwestern Tanzania. PLOS ONE 8: e72753.

64. Niedźwiedzki, G., Brusatte, S. L. & BUTLER, R. J. 2013. Prorotodactylus and Rotodactylus tracks: an ichnological record of dinosauromorphs from the Early-Middle Triassic of Poland. In: Anatomy, Phylogeny and Palaeobiology of Early Archosaurs and their Kin, Nesbitt, S. J., Desojo, J. B. & Irmis, R. B. (eds). Geological Society, London, Special Publications 379: 319-351.

63. Stocker, M. R. & BUTLER, R. J. 2013. Phytosauria. In: Anatomy, Phylogeny and Palaeobiology of Early Archosaurs and their Kin, Nesbitt, S. J., Desojo, J. B. & Irmis, R. B. (eds). Geological Society, London, Special Publications 379: 91-117.

62. Sookias, R. B. & BUTLER, R. J. 2013. Euparkeriidae. In: Anatomy, Phylogeny and Palaeobiology of Early Archosaurs and their Kin, Nesbitt, S. J., Desojo, J. B. & Irmis, R. B. (eds). Geological Society, London, Special Publications 379: 35-48.

61. Ezcurra, M. D., BUTLER, R. J. & Gower, D. J. 2013. Proterosuchia. In: Anatomy, Phylogeny and Palaeobiology of Early Archosaurs and their Kin, Nesbitt, S. J., Desojo, J. B. & Irmis, R. B. (eds). Geological Society, London, Special Publications 379: 9-33.

60. BUTLER, R. J. 2013. “Francosuchustrauthi is not Paleorhinus: implications for Late Triassic vertebrate biostratigraphy. Journal of Vertebrate Paleontology 33: 858–864.

59.
Toljagic, O. &
BUTLER, R. J. 2013. Triassic/Jurassic mass extinction as trigger for the Mesozoic radiation of crocodylomorphs. Biology Letters 9: 20130095.

58. Benson, R. B. J., Mannion, P. D., BUTLER, R. J., Upchurch, P., Goswami, A. & Evans, S. E. 2013. Cretaceous tetrapod fossil record sampling and faunal turnover: implications for biogeography and the rise of modern clades. Palaeogeography, Palaeoclimatology, Palaeoecology 372: 88-107.

57. BUTLER, R. J., Benson, R. B. J. & Barrett, P. M. 2013. Pterosaur diversity: untangling the influences of sampling biases, Lagerstätten, and genuine biodiversity signals. Palaeogeography, Palaeoclimatology, Palaeoecology 372: 78-87.

56. Nesbitt, S. J. & BUTLER, R. J. 2013. Redescription of the archosaur Parringtonia gracilis from the Middle Triassic Manda Beds of Tanzania, and the antiquity of Erpetosuchidae. Geological Magazine 150: 225–238.

55. BUTLER, R. J., Yates, A. M., Rauhut, O. W. M. & Foth, C. 2013. A pathological tail in a basal sauropodomorph dinosaur from South Africa: evidence of traumatic amputation? Journal of Vertebrate Paleontology 33: 224–228.

54. Brusatte, S. L., BUTLER, R. J., Niedźwiedzki, G., Sulej, T., Bronowicz, R. & Satkunas, J. 2013. First record of Mesozoic terrestrial vertebrates from Lithuania: phytosaurs (Diapsida: Archosauriformes) of probable Late Triassic age, with a review of phytosaur biogeography. Geological Magazine 150: 110–122.


2012


53. Han, F.-L., Barrett, P. M., BUTLER, R. J. & Xu, X. 2012. Postcranial anatomy of Jeholosaurus shangyuanensis (Dinosauria, Ornithischia) from the Lower Cretaceous Yixian Formation of China. Journal of Vertebrate Paleontology 32: 1370–1395.

52. Osi, A., Prondvai, E., BUTLER, R. J. & Weishampel, D. B. 2012. Phylogeny, histology and inferred body size evolution in a new rhabdodontid dinosaur from the Late Cretaceous of Hungary. PLoS ONE 7(9): e44318.

51. Mannion, P. D., Benson, R. B. J., Upchurch, P., BUTLER, R. J., Carrano, M. T. & Barrett, P. M. 2012. A temperate palaeodiversity peak in Mesozoic dinosaurs and evidence for Late Cretaceous geographical partitioning. Global Ecology and Biogeography 21: 898–908.

50. Sookias, R. B, Benson, R. B. J. & BUTLER, R. J. 2012. Biology, not environment, drives major patterns in maximum tetrapod body size through time. Biology Letters 8: 674–677.
News coverage: Nature.

49. Brusatte, S. L., BUTLER, R. J., Prieto-Marquez, A. & Norell, M. A. 2012. Dinosaur morphological diversity and the end Cretaceous extinction. Nature Communications 3, article 804.
News & blog coverage: New York Times, N24, Livescience, Telegraph, Smithsonian Dinosaur Tracking blog, Spiegel online, Discovery News, the Guardian, Time, Stern
German language press release: here
AMNH press release and video: here.
Coverage on the LMU homepage: here.

48.
Foth, C., Brusatte, S. L. & BUTLER, R. J. 2012. Do different disparity proxies converge on a common signal? Insights from the cranial morphometrics and evolutionary history of Pterosauria (Diapsida: Archosauria). Journal of Evolutionary Biology 25: 904–915

47. Sookias, R. B., 
BUTLER, R. J.
& Benson, R. B. J.
2012.  Rise of dinosaurs reveals major body size transitions are driven by passive processes of trait evolution. Proceedings of the Royal Society B 279: 2180–2187.
Media & blog coverage: Science, Cosmos, Chinleana 

46. BUTLER, R. J., Porro, L. B., Galton, P. M. & Chiappe, L. M. 2012. Anatomy and cranial functional morphology of the small-bodied dinosaur Fruitadens haagarorum (Ornithischia: Heterodontosauridae) from the Upper Jurassic of the USA. PLoS ONE 7: e31556 (31 pages).
Blog coverage of this paper: PLoS ONE blog, Smithsonian Dinosaur Tracking blog.

45. BUTLER, R. J., Barrett, P. M. & Gower, D. J. 2012. Reassessment of the evidence for postcranial skeletal pneumaticity in Triassic archosaurs, and the early evolution of the avian respiratory system. PLoS ONE 7: e34094 (23 pages).

44. BUTLER, R. J., Brusatte, S. L., Andres, B. & Benson, R. B. J. 2012. How do geological sampling biases affect studies of morphological evolution in deep time? A case study of the Pterosauria (Reptilia: Archosauria). Evolution 66: 147-162.

43. Benson, R. B. J., BUTLER, R. J., Carrano, M. T. & O'Connor, P. M. 2012. Air-filled postcranial bones in theropod dinosaurs: physiological implications and the 'reptile'-bird transition. Biological Reviews 87: 168-193.  


2011

42. BUTLER, R. J., Brusatte, S. L., Reich, M., Nesbitt, S. J., Schoch, R. R. & Hornung, J. J. 2011. The sail-backed reptile Ctenosauriscus from the latest Early Triassic of Germany and the timing and biogeography of the early archosaur radiation. PLoS ONE 6: e25693.
Blog coverage of this paper: Wired Science.

41. Barrett, P. M., BUTLER, R. J., Twitchett, R. J. & Hutt, S. 2011. New material of Valdosaurus canaliculatus (Ornithischia: Ornithopoda) from the Lower Cretaceous of southern England. Special Papers in Palaeontology 86: 131-163.

40. Upchurch, P., Mannion, P. D., Benson, R. B. J., BUTLER, R. J. & Carrano, M. T. 2011. Geological and anthropogenic controls on the sampling of the terrestrial fossil record: a case study from the Dinosauria. In: Comparing the Geological and Fossil Records: Implications for Biodiversity Studies, McGowan, A. J. and Smith, A. B. (eds). Geological Society, London, Special Publication 358: 209-240.

39. Benson, R. B. J. & BUTLER, R. J. 2011. Uncovering the diversification history of marine tetrapods: ecology influences the effect of geological sampling biases. In: Comparing the Geological and Fossil Records: Implications for Biodiversity Studies, McGowan, A. J. and Smith, A. B. (eds.). Geological Society, London, Special Publication 358: 191-207.

38. Norman, D. B., Crompton, A. W., BUTLER, R. J., Porro, L. B. & Charig, A. C. 2011. The Lower Jurassic ornithischian dinosaur Heterodontosaurus tucki Crompton and Charig, 1962: cranial anatomy, functional morphology, taxonomy and relationships. Zoological Journal of the Linnean Society 163: 182-276

37. Porro, L. B., BUTLER, R. J., Barrett, P. M., Moore-Fay, S. & Abel, R. L. 2011 (for 2010). New heterodontosaurid specimens from the Lower Jurassic of southern Africa and the early ornithischian dinosaur radiation. Earth and Environmental Science Transactions of the Royal Society of Edinburgh 101: 351-366.

36. Barrett, P. M., BUTLER, R. J. & Nesbitt, S. J. 2011 (for 2010). The roles of herbivory and omnivory in early dinosaur evolution. Earth and Environmental Science Transactions of the Royal Society of Edinburgh 101: 383-396.

35. BUTLER, R. J., Jin, L., Chen, J. & Godefroit, P. 2011. The postcranial osteology and phylogenetic position of the small ornithischian dinosaur Changchunsaurus parvus from the Quantou Formation (Cretaceous: Aptian-Cenomanian) of Jilin Province, northeastern China. Palaeontology 54: 667-683.

34. BUTLER, R. J., Benson, R. B. J., Carrano, M. T., Mannion, P. D. & Upchurch, P. 2011. Sea-level, dinosaur diversity, and sampling biases: investigating the ‘common cause’ hypothesis in the terrestrial realm. Proceedings of the Royal Society of London B 278: 1165-1170.

33. Brusatte, S. L., Niedźwiedzki, G. & BUTLER, R. J. 2011. Footprints pull origin and diversification of dinosaur stem-lineage deep into Early Triassic. Proceedings of the Royal Society of London B 278: 1107-1113.

32. Wang, X., Pan, R., BUTLER, R. J. & Barrett, P. M. 2011 (for 2010). The postcranial skeleton of the iguanodontian ornithopod Jinzhousaurus yangi from the Lower Cretaceous Yixian Formation of western Liaoning, China. Earth and Environmental Science Transactions of the Royal Society of Edinburgh 101: 135-159.


2010

31. BUTLER, R. J. 2010. The anatomy of the basal ornithischian dinosaur Eocursor parvus from the lower Elliot Formation (Late Triassic) of South Africa. Zoological Journal of the Linnean Society 160: 648-684.

30. Brusatte, S. L., Nesbitt, S. J., Irmis, R. B., BUTLER, R. J., Benton, M. J. & Norell, M. A. 2010. The origin and early radiation of dinosaurs. Earth-Science Reviews 101: 68-100. PDF

29. Osi, A., BUTLER, R. J. & Weishampel, D. B. 2010. A Late Cretaceous ceratopsian dinosaur from Europe with Asian affinities. Nature 465: 466-468. PDF [Ajkaceratops kozmai]
Read a Nature News & Views commentary by Xu Xing on this discovery here. This paper was covered by the BBC, Scientific American, and Science, amongst others.

28. BUTLER, R. J., Barrett, P. M., Kenrick, P. & Penn, M. G. 2010. Testing coevolutionary hypotheses over geological timescales: interactions between Cretaceous dinosaurs and plants. Biological Journal of the Linnean Society 100: 1–15.

27. Benson, R. B. J., BUTLER, R. J., Lindgren, J. & Smith, A. S. 2010. Mesozoic marine tetrapod diversity: mass extinctions and temporal heterogeneity in geologic megabiases affecting vertebrates. Proceedings of the Royal Society of London B 277: 829-834. PDF

26. Jin, L., Chen, J., Zan, S., BUTLER, R. J. & Godefroit, P. 2010. Cranial anatomy of the small ornithischian dinosaur Changchunsaurus parvus from the Quantou Formation (Cretaceous: Aptian–Cenomanian) of Jilin Province, northeastern China. Journal of Vertebrate Paleontology 30: 196-214.

25. BUTLER, R. J., Galton, P. M., Porro, L. B., Chiappe, L. M., Henderson, D. M. & Erickson, G. M. 2010. Lower limits of ornithischian dinosaur body size inferred from a new Upper Jurassic heterodontosaurid from North America. Proceedings of the Royal Society of London B 277: 375-381. PDF [Fruitadens haagarorum]
This paper was covered by the BBC, National Geographic, and the New York Times, amongst others.


2009

24. BUTLER, R. J., Barrett, P. M., Abel, R. L. & Gower, D. J. 2009. A possible ctenosauriscid archosaur from the Middle Triassic Manda Beds of Tanzania. Journal of Vertebrate Paleontology 29: 1022-1031

23. BUTLER, R. J., Barrett, P. M. & Gower, D. J. 2009. Postcranial skeletal pneumaticity and air-sacs in the earliest pterosaurs. Biology Letters 5: 557-560. PDF

22. BUTLER, R. J., Barrett, P. M., Nowbath, S. & Upchurch, P. 2009. Estimating the effects of the rock record on pterosaur diversity patterns: implications for hypotheses of bird/pterosaur competitive replacement. Paleobiology 35: 432-446.

21. Brusatte, S. L., BUTLER. R. J., Sulej, T. & Niedzwiedzki, G. 2009. The taxonomy and anatomy of rauisuchian archosaurs from the Late Triassic of Germany and Poland. Acta Palaeontologica Polonica 54: 221-230. PDF   

20. Barrett, P. M., BUTLER, R. J., Wang, X.-L. & Xu, X. 2009. Cranial anatomy of the iguanodontoid ornithopod Jinzhousaurus yangi from the Lower Cretaceous Yixian Formation of China. Acta Palaeontologica Polonica 54: 35-48. PDF

19. BUTLER, R. J. & Sullivan, R. M. 2009. The phylogenetic position of the ornithischian dinosaur Stenopelix valdensis from the Lower Cretaceous of Germany: implications for the early fossil record of Pachycephalosauria. Acta Palaeontologica Polonica 54: 21-34. PDF

18. BUTLER, R. J., Barrett, P. M., Kenrick, P. & Penn, M. G. 2009. Diversity patterns amongst dinosaurs and plants during the Cretaceous: implications for hypotheses of dinosaur/angiosperm co-evolution. Journal of Evolutionary Biology 22: 446-459. PDF

17. BUTLER, R. J., Barrett, P. M., Kenrick, P. & Penn, M. G. 2009. Testing co-evolutionary hypotheses over geological timescales: interactions between Mesozoic non-avian dinosaurs and cycads. Biological Reviews 84: 73-89.

16. BUTLER, R. J. & Zhao, Q. 2009. The small-bodied ornithischian dinosaurs Micropachycephalosaurus hongtuyanensis and Wannanosaurus yansiensis from the Late Cretaceous of China. Cretaceous Research 30: 63-77.

 

2008

15. Barrett, P. M., BUTLER, R. J., Edwards, N. P. & Milner, A. R. 2008. Pterosaur distribution in time and space: an atlas. Zitteliana B 28: 61-107. PDF

14. BUTLER, R. J., Upchurch, P. & Norman, D. B. 2008. The phylogeny of the ornithischian dinosaurs. Journal of Systematic Palaeontology 6: 1–40.

13. BUTLER, R. J., Porro, L. B. & Norman, D. B. 2008. A juvenile skull of the primitive ornithischian Heterodontosaurus tucki from the ‘Stormberg’ of South Africa. Journal of Vertebrate Paleontology 28: 702-711.

12. BUTLER, R. J. & Goswami, A. 2008. Body size evolution in Mesozoic birds: little evidence for Cope's rule. Journal of Evolutionary Biology 26: 1673-1682. PDF

11. BUTLER, R. J. & Barrett, P. M. 2008. Palaeoenvironmental controls on the distribution of Cretaceous herbivorous dinosaurs. Naturwissenschaften 95: 1027-1032

10. BUTLER, R. J. & Galton, P. M. 2008. The ‘dermal armour’ of the Wealden dinosaur Hypsilophodon: a reappraisal. Cretaceous Research 29: 636-642.

9. Chen, J., BUTLER , R. J. & Jin, L. 2008. New material of large-bodied ornithischian dinosaurs, including an iguanodontian ornithopod, from the Quantou Formation (middle Cretaceous: Aptian–Cenomanian) of Jilin Province, northeastern China. Neues Jahrbuch für Geologie und Paläontologie, Abhandlungen 248: 309-314.

8. Barrett, P. M, BUTLER, R. J., Novas, F. E., Moore-Fay, S., Moody, J. M., Clark, J. M. & Sanchez-Villagra, M. R. 2008. Dinosaur remains from the La Quinta Formation (Lower or Middle Jurassic) of the Venezuelan Andes. Paläontologische Zeitschrift 82: 163–177.

 

2007

7. BUTLER, R. J., Smith, R. M. H. & Norman, D. B. 2007. A primitive ornithischian dinosaur from the Late Triassic of South Africa, and the early evolution and diversification of Ornithischia. Proceedings of the Royal Society of London B 274: 2041–2046. PDF

6. Norman, D. B., BUTLER, R. J. & Maidment, S. C. R. 2007. Reconsidering the status and affinities of the ornithischian dinosaur Tatisaurus oehleri Simmons, 1965. Zoological Journal of the Linnean Society 150: 865–874. PDF

5. BUTLER, R. J. & Upchurch, P. 2007. The problem of highly incomplete taxa and the phylogenetic relationships of the theropod dinosaur Juravenator starki. Journal of Vertebrate Paleontology 27: 253–256


2006 

4. Corfe, I. J. & BUTLER, R. J. 2006. Comment on ‘A Well-Preserved Archaeopteryx Specimen with Theropod Features’. Science 313:1238. (Published online; doi:10.1126/science.1130800)

3. BUTLER, R. J., Porro, L. B. & Heckert, A. B. 2006. A supposed heterodontosaurid tooth from the Rhaetian of Switzerland and a reassessment of the European Late Triassic record of Ornithischia (Dinosauria). Neues Jahrbuch für Geologie und Paläontologie, Monatshefte 10: 613–633. 


2005

2. Barrett, P. M., BUTLER, R. J. & Knoll, F. 2005. Small-bodied ornithischian dinosaurs from the Middle Jurassic of Sichuan, China. Journal of Vertebrate Paleontology 25: 823–834

1. BUTLER, R. J. 2005. The ‘fabrosaurid’ ornithischian dinosaurs of the Upper Elliot Formation (Lower Jurassic) of South Africa and Lesotho. Zoological Journal of the Linnean Society, 145: 175–218. PDF


Conference papers

3. BUTLER, R. J., Barrett, P. M., Kenrick, P. & Penn, M. G. 2008. Testing co-evolutionary hypotheses over geological timescales using GIS; pp 214–216 in D. Lambrick (ed.), Proceedings of the GIS Research UK 16th Annual Conference, GISRUK 2008, Manchester Metropolitan University, 2–4 April 2008

2. BUTLER, R. J., Upchurch, P., Norman, D. B., & Parish, J. C. 2006. A biogeographical analysis of the ornithischian dinosaurs; pp. 13–16 & 151 in P.M. Barrett and S. E. Evans (eds.), Ninth International Symposium on Mesozoic Terrestrial Ecosystems and Biota, Abstracts and Proceedings. The Natural History Museum, London. [Short paper and abstract]

1. Evans, S. E., Barrett, P. M., Hilton, J., BUTLER, R. J., Jones, M., Liang, M. M., Parish, J. C., Rayfield, E., Sigogneau-Russell, D. & Underwood, C. J. 2006. The Middle Jurassic vertebrate assemblage of Skye, Scotland; pp. 36–39 & 157 in P.M. Barrett and S. E. Evans (eds.), Ninth International Symposium on Mesozoic Terrestrial Ecosystems and Biota, Abstracts and Proceedings. The Natural History Museum, London. [Short paper and abstract]