Publications

This page lists publications produced by all members of the group during (or resulting from) their time within the Archosauromorph Research Group during its time at the University of Birmingham (from August 2013) and the LMU München (September 2011–July 2013). Publications for Richard Butler resulting from work carried out prior to the foundation of the group (with BSPG, NHM or Cambridge affiliations) are also listed. Group members are highlighted in bold and capitals. Details of individual researchers' publications are also present on their homepages

Below: Slideshow of recent (2012–2014) papers published by our group.

In press

EZCURRA, M. D. & BUTLER, R. J. Taxonomy of the proterosuchid archosauriforms (Diapsida: Archosauromorpha) from the earliest Triassic of South Africa, and implications for the early archosauriform radiation. Palaeontology.

Brusatte, S. L., 
BUTLER, R. J., Mateus, O. & Steyer, J. S. A new species of Metoposaurus from the Late Triassic of Portugal and comments on the systematics and biogeography of metoposaurid temnospondyls. Journal of Vertebrate Paleontology.

2014

Upchurch, P., Andres, B., BUTLER, R. J. & Barrett, P. M. (published online). An analysis of pterosaurian biogeography: implications for the evolutionary history and fossil record quality of the first flying vertebrates. Historical Biology.

Brusatte, S. L., BUTLER, R. J., Barrett, P. M., Carrano, M. T., Evans, D. C., Lloyd, G. T., Mannion, P. D., Norell, M. A., Peppe, D. J., Upchurch, P. & Williamson, T. E. (published online). The extinction of the dinosaurs. Biological Reviews.


SOOKIAS, R. B., Sennikov, A. G., Gower, D. J. & BUTLER, R. J. (published online). The monophyly of Euparkeriidae (Reptilia: Archosauriformes) and the origins of crown Archosauria: a revision of Dorosuchus neoetus from the Middle Triassic of Russia. Palaeontology.

Niedźwiedzki
, G., Brusatte, S. L., Sulej, T. & BUTLER, R. J. (published online). Basal dinosauriform and theropod dinosaurs from the middle-late Norian (Late Triassic) of Poland: implications for Triassic dinosaur evolution and distribution. Palaeontology.

EZCURRA, M. D., Trotteyn, M. J., Fiorelli, L. E., Taborda, J. R. A, Baczko, von, M. B., Iberlucea, M., Desojo, J.B. (published online). The oldest rhynchosaur from Argentina: a Middle Triassic rhynchosaurid from the Chañares Formation. Paläontologische Zeitschrift. 

Barrett, P. M., BUTLER, R. J., Mundil, R., Scheyer, T. M., Irmis, R. B. & Sánchez-Villagra, M. R. 2014. A palaeoequatorial ornithischian and new constraints on early dinosaur diversification. Proceedings of the Royal Society B 281: 20141147

Mateus, O., BUTLER, R. J., Brusatte, S. L., Steyer, J. S. & Whiteside, J. H. 2014. The first phytosaur (Diapsida, Archosauriformes) from the Late Triassic of the Iberian Peninsula. Journal of Vertebrate Paleontology 34:970-975.

Porfiri, J. D., Novas, F. E., Calvo, J. O., Agnolín, F. L., EZCURRA, M. D. & Cerda, I. A. 2014. Juvenile specimen of Megaraptor (Dinosauria, Theropoda) sheds light about tyrannosauroid radiation. Cretaceous Research 51: 35-55.

BUTLER, R. J., Sullivan, C., EZCURRA, M. D., Liu, J., Lecuona, A. & SOOKIAS, R. B. 2014. New clade of enigmatic early archosaurs yields insights into early pseudosuchian phylogeny and the biogeography of the archosaur radiation. BMC Evolutionary Biology 14:128. 

Benson, R. B. J., Frigot, R. A., Goswami, A., Andres, B. & BUTLER, R. J. 2014. Competition and constraint drove Cope's rule in the evolution of giant flying reptiles. Nature Communications 5:3567.

Delsate, D. & EZCURRA, M. D. 2014. The first Early Jurassic (late Hettangian) theropod dinosaur remains from the Grand Duchy of Luxembourg. Geologica Belgica 17: 175-181.

EZCURRA, M. D.
, Scheyer, T. & BUTLER, R. J. 2014. The origin and early evolution of Sauria: reassessing the Permian saurian fossil record and the timing of the crocodile-lizard divergence. PLOS ONE 9:
e89165.

BUTLER, R. J.
, Rauhut, O. W. M., Stocker, M. R. & Bronowicz, R. 2014. Redescription of the phytosaurs Paleorhinus ("Francosuchus") angustifrons and Ebrachosuchus neukami from Germany, with implications for Late Triassic biochronology. Zoological Journal of the Linnean Society 170: 155–208.

EZCURRA, M. D. 2014. The osteology of the basal archosauromorph Tasmaniosaurus triassicus from the Lower Triassic of Tasmania, Australia. PLOS ONE 9: e86864.

2013

Fiorelli, L. E., EZCURRA, M. D., Hechenleitner, E. M., Arganaraz, E., Taborda, J. R. A., Trotteyn, M. J., von Baczko, M. B. & Desojo, J. B. 2013. The oldest known communal latrines provide evidence of gregarism in Triassic megaherbivores. Scientific Reports 3: 3348 (DOI: 10.1038/srep03348)

Novas, F. E., Agnolín, F. L., EZCURRA, M. D., Porfiri, J. & Canale, J. I. 2013. Evolution of the carnivorous dinosaurs during the Cretaceous: the evidence from Patagonia. Cretaceous Research 45: 174-215.


Nesbitt, S. J., BUTLER, R. J. & Gower, D. J. 2013. A new archosauriform (Reptilia: Diapsida) from the Manda beds (Middle Triassic) of southwestern Tanzania. PLOS ONE 8: e72753.

SOOKIAS, R. B., Budd, G. E. & Kear, B. P. 2013. Mesozoic fossil sustainability: synoptic case studies of resource management. GFF 135: 131-143.

Niedźwiedzki, G., Brusatte, S. L. & BUTLER, R. J. 2013.
Prorotodactylus and Rotodactylus tracks: an ichnological record of dinosauromorphs from the Early-Middle Triassic of Poland.
In: Anatomy, Phylogeny and Palaeobiology of Early Archosaurs and their Kin, Nesbitt, S. J., Desojo, J. B. & Irmis, R. B. (eds). Geological Society, London, Special Publications 379: 319-351.

Baczko, M. B., von & EZCURRA, M. D. 2013. Ornithosuchidae. In: Anatomy, Phylogeny and Palaeobiology of Early Archosaurs and their Kin, Nesbitt, S. J., Desojo, J. B. & Irmis, R. B. (eds). Geological Society, London, Special Publications 379: 187-202.

Stocker, M. R. & BUTLER, R. J. 2013. Phytosauria. In: Anatomy, Phylogeny and Palaeobiology of Early Archosaurs and their Kin, Nesbitt, S. J., Desojo, J. B. & Irmis, R. B. (eds). Geological Society, London, Special Publications 379: 91-117.

Sues, H.-D., Desojo, J. B. & EZCURRA, M. D. 2013. Doswelliidae. In: Anatomy, Phylogeny and Palaeobiology of Early Archosaurs and their Kin, Nesbitt, S. J., Desojo, J. B. & Irmis, R. B. (eds). Geological Society, London, Special Publications 379: 49-58.

SOOKIAS, R. B. & BUTLER, R. J. 2013. Euparkeriidae. In: Anatomy, Phylogeny and Palaeobiology of Early Archosaurs and their Kin, Nesbitt, S. J., Desojo, J. B. & Irmis, R. B. (eds). Geological Society, London, Special Publications 379: 35-48.

EZCURRA, M. D., BUTLER, R. J. & Gower, D. J. 2013. Proterosuchia. In: Anatomy, Phylogeny and Palaeobiology of Early Archosaurs and their Kin, Nesbitt, S. J., Desojo, J. B. & Irmis, R. B. (eds). Geological Society, London, Special Publications 379: 9-33.

BUTLER, R. J.
2013. “Francosuchustrauthi is not Paleorhinus: implications for Late Triassic vertebrate biostratigraphy. Journal of Vertebrate Paleontology 33: 858–864.

TOLJAGIC, O.
& BUTLER, R. J. 2013. Triassic/Jurassic mass extinction as trigger for the Mesozoic radiation of crocodylomorphs. Biology Letters 9: 20130095.

Benson, R. B. J., Mannion, P. D., BUTLER, R. J., Upchurch, P., Goswami, A. & Evans, S. E. 2013. Cretaceous tetrapod fossil record sampling and faunal turnover: implications for biogeography and the rise of modern clades. Palaeogeography, Palaeoclimatology, Palaeoecology 372: 88-107.

BUTLER, R. J., Benson, R. B. J. & Barrett, P. M. 2013. Pterosaur diversity: untangling the influences of sampling biases, Lagerstätten, and genuine biodiversity signals. Palaeogeography, Palaeoclimatology, Palaeoecology 372: 78-87.

Nesbitt, S. J. & BUTLER, R. J. 2013. Redescription of the archosaur Parringtonia gracilis from the Middle Triassic Manda Beds of Tanzania, and the antiquity of Erpetosuchidae. Geological Magazine 150: 225–238.

BUTLER, R. J.
, Yates, A. M., Rauhut, O. W. M. & Foth, C. 2013. A pathological tail in a basal sauropodomorph dinosaur from South Africa: evidence of traumatic amputation? Journal of Vertebrate Paleontology 33: 224–228.

Brusatte, S. L., BUTLER, R. J., Niedźwiedzki, G., Sulej, T., Bronowicz, R. & Satkunas, J. 2013. First record of Mesozoic terrestrial vertebrates from Lithuania: phytosaurs (Diapsida: Archosauriformes) of probable Late Triassic age, with a review of phytosaur biogeography. Geological Magazine 150: 110–122.


2012

Novas, F. E., Kundrat, M., Agnolín, F. L., EZCURRA, M. D., Ahlberg, P. E., Isasi, M. P., Arriagada, A. & Chafrat, P. 2012. A new large pterosaur from the Late Cretaceous of Patagonia. Journal of Vertebrate Paleontology 32: 1147-1452.

Han, F.-L., Barrett, P. M., BUTLER, R. J. & Xu, X. 2012. Postcranial anatomy of Jeholosaurus shangyuanensis (Dinosauria, Ornithischia) from the Lower Cretaceous Yixian Formation of China. Journal of Vertebrate Paleontology 32: 1370–1395.

Osi, A., Prondvai, E., BUTLER, R. J. & Weishampel, D. B. 2012. Phylogeny, histology and inferred body size evolution in a new rhabdodontid dinosaur from the Late Cretaceous of Hungary. PLoS ONE 7(9): e44318.

Novas, F. E., EZCURRA, M. D., Agnolín, F. L., Pol, D. & Ortíz, R. 2012. New Patagonian Cretaceous theropod sheds light about the early radiation of Coelurosauria.
Revista del Museo Argentino de Ciencias Naturales, nueva serie, 14: 57–81.

EZCURRA, M. D. 2012. Comments on the taxonomic diversity and paleobiogeography of the earliest known dinosaur assemblages (late Carnian-earliest Norian). Historia Natural, tercera serie 2: 49–71.

SOOKIAS, R. B, Benson, R. B. J. & BUTLER, R. J. 2012. Biology, not environment, drives major patterns in maximum tetrapod body size through time. Biology Letters 8: 674–677.
News coverage: Nature.

Mannion, P. D., Benson, R. B. J., Upchurch, P., BUTLER, R. J., Carrano, M. T. & Barrett, P. M. 2012. A temperate palaeodiversity peak in Mesozoic dinosaurs and evidence for Late Cretaceous geographical partitioning. Global Ecology and Biogeography 21: 898–908.

EZCURRA, M. D. & Agnolín, F. L. 2012. An abelisauroid dinosaur from the Middle Jurassic of Laurasia and its implications on theropod palaeobiogeography and evolution. Proceedings of the Geologists’ Association 123: 500–507.

Brusatte, S. L., BUTLER, R. J., Prieto-Marquez, A. & Norell, M. A. 2012. Dinosaur morphological diversity and the end Cretaceous extinction. Nature Communications 3, article 804.

News & blog coverage: New York Times, N24, Livescience, Telegraph, Smithsonian Dinosaur Tracking blog, Spiegel online, Discovery News, the Guardian, Time, Stern
German language press release: here.  

AMNH press release and video: here.
Coverage on the LMU homepage: here.

Foth, C., Brusatte, S. L. & BUTLER, R. J. 2012. Do different disparity proxies converge on a common signal? Insights from the cranial morphometrics and evolutionary history of Pterosauria (Diapsida: Archosauria). Journal of Evolutionary Biology 25: 904–915

SOOKIAS, R. B.BUTLER, R. J. & Benson, R. B. J. 2012. Rise of dinosaurs reveals major body size transitions are driven by passive processes of trait evolution. Proceedings of the Royal Society B 279: 2180–2187.
Media & blog coverage: Science, Cosmos, Chinleana 

BUTLER, R. J., Porro, L. B., Galton, P. M. & Chiappe, L. M. 2012. Anatomy and cranial functional morphology of the small-bodied dinosaur Fruitadens haagarorum (Ornithischia: Heterodontosauridae) from the Upper Jurassic of the USA. PLoS ONE 7: e31556 (31 pages).
Blog coverage of this paper: PLoS ONE blog, Smithsonian Dinosaur Tracking blog.

BUTLER, R. J., Barrett, P. M. & Gower, D. J. 2012. Reassessment of the evidence for postcranial skeletal pneumaticity in Triassic archosaurs, and the early evolution of the avian respiratory system. PLoS ONE 7: e34094 (23 pages).

BUTLER, R. J., Brusatte, S. L., Andres, B. & Benson, R. B. J. 2012. How do geological sampling biases affect studies of morphological evolution in deep time? A case study of the Pterosauria (Reptilia: Archosauria). Evolution 66: 147-162.

Benson, R. B. J., BUTLER, R. J., Carrano, M. T. & O'Connor, P. M. 2012. Air-filled postcranial bones in theropod dinosaurs: physiological implications and the 'reptile'-bird transition. Biological Reviews 87: 168-193.


2011



BUTLER, R. J., Brusatte, S. L., Reich, M., Nesbitt, S. J., Schoch, R. R. & Hornung, J. J. 2011. The sail-backed reptile Ctenosauriscus from the latest Early Triassic of Germany and the timing and biogeography of the early archosaur radiation. PLoS ONE 6: e25693 (28 pages).
Blog coverage of this paper: Wired Science.

Barrett, P. M., BUTLER, R. J., Twitchett, R. J. & Hutt, S. 2011. New material of Valdosaurus canaliculatus (Ornithischia: Ornithopoda) from the Lower Cretaceous of southern England. Special Papers in Palaeontology 86: 131-163.

Upchurch, P., Mannion, P. D., Benson, R. B. J., BUTLER, R. J. & Carrano, M. T. 2011. Geological and anthropogenic controls on the sampling of the terrestrial fossil record: a case study from the Dinosauria. In: Comparing the Geological and Fossil Records: Implications for Biodiversity Studies, McGowan, A. J. and Smith, A. B. (eds). Geological Society, London, Special Publication 358: 209-240.

Benson, R. B. J. & BUTLER, R. J. 2011. Uncovering the diversification history of marine tetrapods: ecology influences the effect of geological sampling biases. In: Comparing the Geological and Fossil Records: Implications for Biodiversity Studies, McGowan, A. J. and Smith, A. B. (eds.). Geological Society, London, Special Publication 358: 191-207.

Norman, D. B., Crompton, A. W., BUTLER, R. J., Porro, L. B. & Charig, A. C. 2011. The Lower Jurassic ornithischian dinosaur Heterodontosaurus tucki Crompton and Charig, 1962: cranial anatomy, functional morphology, taxonomy and relationships. Zoological Journal of the Linnean Society 163: 182-276

Porro, L. B., BUTLER, R. J., Barrett, P. M., Moore-Fay, S. & Abel, R. L. 2011 (for 2010). New heterodontosaurid specimens from the Lower Jurassic of southern Africa and the early ornithischian dinosaur radiation. Earth and Environmental Science Transactions of the Royal Society of Edinburgh 101: 351-366.

Barrett, P. M., BUTLER, R. J. & Nesbitt, S. J. 2011 (for 2010). The roles of herbivory and omnivory in early dinosaur evolution. Earth and Environmental Science Transactions of the Royal Society of Edinburgh 101: 383-396.

BUTLER, R. J., Jin, L., Chen, J. & Godefroit, P. 2011. The postcranial osteology and phylogenetic position of the small ornithischian dinosaur Changchunsaurus parvus from the Quantou Formation (Cretaceous: Aptian-Cenomanian) of Jilin Province, northeastern China. Palaeontology 54: 667-683.

BUTLER, R. J., Benson, R. B. J., Carrano, M. T., Mannion, P. D. & Upchurch, P. 2011. Sea-level, dinosaur diversity, and sampling biases: investigating the ‘common cause’ hypothesis in the terrestrial realm. Proceedings of the Royal Society of London B 278: 1165-1170.

Brusatte, S. L., Niedźwiedzki, G. & BUTLER, R. J. 2011. Footprints pull origin and diversification of dinosaur stem-lineage deep into Early Triassic. Proceedings of the Royal Society of London B 278: 1107-1113.

Wang, X., Pan, R., BUTLER, R. J. & Barrett, P. M. 2011 (for 2010). The postcranial skeleton of the iguanodontian ornithopod Jinzhousaurus yangi from the Lower Cretaceous Yixian Formation of western Liaoning, China. Earth and Environmental Science Transactions of the Royal Society of Edinburgh 101: 135-159.

2010

BUTLER, R. J. 2010. The anatomy of the basal ornithischian dinosaur Eocursor parvus from the lower Elliot Formation (Late Triassic) of South Africa. Zoological Journal of the Linnean Society 160: 648-684.

Brusatte, S. L., Nesbitt, S. J., Irmis, R. B., BUTLER, R. J., Benton, M. J. & Norell, M. A. 2010. The origin and early radiation of dinosaurs. Earth-Science Reviews 101: 68-100. PDF

Osi, A., BUTLER, R. J. & Weishampel, D. B. 2010. A Late Cretaceous ceratopsian dinosaur from Europe with Asian affinities. Nature 465: 466-468. PDF [Ajkaceratops kozmai]
Read a Nature News & Views commentary by Xu Xing on this discovery here. This paper was covered by the BBC, Scientific American, and Science, amongst others.

BUTLER, R. J., Barrett, P. M., Kenrick, P. & Penn, M. G. 2010. Testing coevolutionary hypotheses over geological timescales: interactions between Cretaceous dinosaurs and plants. Biological Journal of the Linnean Society 100: 1–15.

Benson, R. B. J., BUTLER, R. J., Lindgren, J. & Smith, A. S. 2010. Mesozoic marine tetrapod diversity: mass extinctions and temporal heterogeneity in geologic megabiases affecting vertebrates. Proceedings of the Royal Society of London B 277: 829-834. PDF

Jin, L., Chen, J., Zan, S., BUTLER, R. J. & Godefroit, P. 2010. Cranial anatomy of the small ornithischian dinosaur Changchunsaurus parvus from the Quantou Formation (Cretaceous: Aptian–Cenomanian) of Jilin Province, northeastern China. Journal of Vertebrate Paleontology 30: 196-214.

BUTLER, R. J., Galton, P. M., Porro, L. B., Chiappe, L. M., Henderson, D. M. & Erickson, G. M. 2010. Lower limits of ornithischian dinosaur body size inferred from a new Upper Jurassic heterodontosaurid from North America. Proceedings of the Royal Society of London B 277: 375-381. PDF [Fruitadens haagarorum]
This paper was covered by the BBC, National Geographic, and the New York Times, amongst others.


2009

BUTLER, R. J., Barrett, P. M., Abel, R. L. & Gower, D. J. 2009. A possible ctenosauriscid archosaur from the Middle Triassic Manda Beds of Tanzania. Journal of Vertebrate Paleontology 29: 1022-1031

BUTLER, R. J., Barrett, P. M. & Gower, D. J. 2009. Postcranial skeletal pneumaticity and air-sacs in the earliest pterosaurs. Biology Letters 5: 557-560. PDF

BUTLER, R. J., Barrett, P. M., Nowbath, S. & Upchurch, P. 2009. Estimating the effects of the rock record on pterosaur diversity patterns: implications for hypotheses of bird/pterosaur competitive replacement. Paleobiology 35: 432-446.

Brusatte, S. L., BUTLER. R. J., Sulej, T. & Niedzwiedzki, G. 2009. The taxonomy and anatomy of rauisuchian archosaurs from the Late Triassic of Germany and Poland. Acta Palaeontologica Polonica 54: 221-230. PDF   

Barrett, P. M., BUTLER, R. J., Wang, X.-L. & Xu, X. 2009. Cranial anatomy of the iguanodontoid ornithopod Jinzhousaurus yangi from the Lower Cretaceous Yixian Formation of China. Acta Palaeontologica Polonica 54: 35-48. PDF

BUTLER, R. J. & Sullivan, R. M. 2009. The phylogenetic position of the ornithischian dinosaur Stenopelix valdensis from the Lower Cretaceous of Germany: implications for the early fossil record of Pachycephalosauria. Acta Palaeontologica Polonica 54: 21-34. PDF

BUTLER, R. J., Barrett, P. M., Kenrick, P. & Penn, M. G. 2009. Diversity patterns amongst dinosaurs and plants during the Cretaceous: implications for hypotheses of dinosaur/angiosperm co-evolution. Journal of Evolutionary Biology 22: 446-459. PDF

BUTLER, R. J., Barrett, P. M., Kenrick, P. & Penn, M. G. 2009. Testing co-evolutionary hypotheses over geological timescales: interactions between Mesozoic non-avian dinosaurs and cycads. Biological Reviews 84: 73-89.

BUTLER, R. J. & Zhao, Q. 2009. The small-bodied ornithischian dinosaurs Micropachycephalosaurus hongtuyanensis and Wannanosaurus yansiensis from the Late Cretaceous of China. Cretaceous Research 30: 63-77.

 

2008

Barrett, P. M., BUTLER, R. J., Edwards, N. P. & Milner, A. R. 2008. Pterosaur distribution in time and space: an atlas. Zitteliana B 28: 61-107. PDF

BUTLER, R. J., Upchurch, P. & Norman, D. B. 2008. The phylogeny of the ornithischian dinosaurs. Journal of Systematic Palaeontology 6: 1–40.

BUTLER, R. J., Porro, L. B. & Norman, D. B. 2008. A juvenile skull of the primitive ornithischian Heterodontosaurus tucki from the ‘Stormberg’ of South Africa. Journal of Vertebrate Paleontology 28: 702-711.

BUTLER, R. J. & Goswami, A. 2008. Body size evolution in Mesozoic birds: little evidence for Cope's rule. Journal of Evolutionary Biology 26: 1673-1682. PDF

BUTLER, R. J. & Barrett, P. M. 2008. Palaeoenvironmental controls on the distribution of Cretaceous herbivorous dinosaurs. Naturwissenschaften 95: 1027-1032

BUTLER, R. J. & Galton, P. M. 2008. The ‘dermal armour’ of the Wealden dinosaur Hypsilophodon: a reappraisal. Cretaceous Research 29: 636-642.

Chen, J., BUTLER , R. J. & Jin, L. 2008. New material of large-bodied ornithischian dinosaurs, including an iguanodontian ornithopod, from the Quantou Formation (middle Cretaceous: Aptian–Cenomanian) of Jilin Province, northeastern China. Neues Jahrbuch für Geologie und Paläontologie, Abhandlungen 248: 309-314.

Barrett, P. M, BUTLER, R. J., Novas, F. E., Moore-Fay, S., Moody, J. M., Clark, J. M. & Sanchez-Villagra, M. R. 2008. Dinosaur remains from the La Quinta Formation (Lower or Middle Jurassic) of the Venezuelan Andes. Paläontologische Zeitschrift 82: 163–177.


2005–2007

BUTLER, R. J., Smith, R. M. H. & Norman, D. B. 2007. A primitive ornithischian dinosaur from the Late Triassic of South Africa, and the early evolution and diversification of Ornithischia. Proceedings of the Royal Society of London B 274: 2041–2046. PDF

Norman, D. B., BUTLER, R. J. & Maidment, S. C. R. 2007. Reconsidering the status and affinities of the ornithischian dinosaur Tatisaurus oehleri Simmons, 1965. Zoological Journal of the Linnean Society 150: 865–874. PDF

BUTLER, R. J. & Upchurch, P. 2007. The problem of highly incomplete taxa and the phylogenetic relationships of the theropod dinosaur Juravenator starki. Journal of Vertebrate Paleontology 27: 253–256

Corfe, I. J. & BUTLER, R. J. 2006. Comment on ‘A Well-Preserved Archaeopteryx Specimen with Theropod Features’. Science 313:1238. (Published online; doi:10.1126/science.1130800)

BUTLER, R. J., Porro, L. B. & Heckert, A. B. 2006. A supposed heterodontosaurid tooth from the Rhaetian of Switzerland and a reassessment of the European Late Triassic record of Ornithischia (Dinosauria). Neues Jahrbuch für Geologie und Paläontologie, Monatshefte 10: 613–633. 

Barrett, P. M., BUTLER, R. J. & Knoll, F. 2005. Small-bodied ornithischian dinosaurs from the Middle Jurassic of Sichuan, China. Journal of Vertebrate Paleontology 25: 823–834

BUTLER, R. J. 2005. The ‘fabrosaurid’ ornithischian dinosaurs of the Upper Elliot Formation (Lower Jurassic) of South Africa and Lesotho. Zoological Journal of the Linnean Society, 145: 175–218. PDF